RhinobatosRajaHeliobatisAsterotrygonHexatrygonPlesiobatisUrolophusTrygonopteraUrobatisUrotrygonParatrygonPlesiotrygonPotamotrygonHimanturaATaeniuraHimanturaBDasyatisPteroplatytrygonGymnuraMyliobatisAetobatusRhinopteraMobulaOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.de_Carvalho_etal_2004ab