DromaeosauridaeArchaeopteryxJeholornisChongmingiaSapeornisConfuciusornis_sanctusChangchengornis_hengdaoziensisEoconfuciusornis_zhengiConfuciusornis_duiBoluochiaConcornisElsornisEoalulavisCathayornisEocathayornisEoenantiornisGobipteryxLongipteryxLongirostravisNeuquenornisPengornisEopengornisProtopteryxRapaxavisShanweiniaoVescornisQilianiaDunhuangiaPiscivorenantiornisLinyiornisSulcavisBohaiornisLongusunguisShenqiornisZhouornisParabohaiornisFortunguavisPterygornisCruralispenniaMonoenantiornisArchaeorhynchusSchizoouraBellulornisVoronaJianchangornisSonglingornisLongicrusavisApsaravisHongshanornisArchaeornithuraParahongshanornisTianyuornisYanornisPatagopteryxYixianornisPiscivoravisIteravisGansusIchthyornisHesperornisParahesperornsiEnaliornisBaptornis_advenusBaptornis_varneriVegavisAnasGallusIVPP_V18929IVPP_V25033IVPP_V24194OsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Wang_etal_2018b