Corvus_moneduloidesAlca_tordaPinguinus_impennisAlle_alleUria_aalgeCepphus_columbaBrachyramphus_marmoratusSynthliboramphus_antiquusAethia_cristatellaFratercula_corniculataMancallinae_spStercorarius_longicaudusSterna_anaethetusRynchops_nigerRissa_tridactylaLarus_argentatusStiltia_isabellaCharadrius_vociferusHalcyornis_toliapicusBubo_virginianusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Smith_et_Clarke_2012a