Colbertia_magellanicaAnisotemnus_distentusThomashuxleya_externaCoquenia_bondiMartinmiguelia_fernandeziTaubatherium_paulacoutoiAncylocoelus_frequensLeontinia_gaudryiElmerriggsia_fieldiaScarrittia_canquelensisAnayatherium_ekecoaAnayatherium_fortisHuilatherium_pluriplicatumColpodon_propinquusColpodon_antucoensisRhynchippus_equinusRhynchippus_pumilusArgyrohippus_fraterculusEurygenium_pacegnumNesodon_imbricatusAdinotherium_ovinumHomalodotherium_cunninghamiFedericoanaya_sallaensisProtypotherium_australeHegetotherium_mirabileTrachytherus_alloxusMesotherium_cristatumOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Shockey_etal_2012a