VictoriapithecusAllenopithecusParapapio_lothagamensisMacaca_spCercocebus_agilisCercocebus_atysCercocebus_chrysogasterCercocebus_torquatusLophocebus_albigenaLophocebus_aterrimusMandrillus_leucophaeusMandrillus_sphinxPapio_anubisPapio_cynocephalusPapio_hamadryasPapio_kindaePapio_papioPapio_ursinusPapio_robinsoniPapio_angusticepsTheropithecus_geladaDinopithecusGorgopithecusSoromandrillusParapapio_adoParapapio_broomiParapapio_jonesiParapapio_whiteiPliopapioProcercocebusTheropithecus_brumptiTheropithecus_oswaldi_dartiTheropithecus_baringensisLophocebus_sp_novPapio_izodiRungwecebus_kipunjiOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Pugh_et_Gilbert_2018a