Larus_dominicanus_Southern_blackbacked_gullAptenodytes_forsteri_Emperor_penguinAptenodytes_patagonicus_King_penguinPygoscelis_antarctica_Chinstrap_penguinPygoscelis_adeliae_Adelie_penguinPygoscelis_papua_Gentoo_penguinSpheniscus_demersus_Black_footed_penguinSpheniscus_magellanicus_Magellanic_penguinEudyptula_minor_Little_blue_penguinMegadyptes_antipodes_Yellow_eyed_penguinEudyptes_pachyrhynchus_Fiordland_penguinEudyptes_robustus_Snares_penguinEudyptes_sclateri_Erect_crested_penguinEudyptes_chrysocome_Rockhopper_penguinEudyptes_chrysolophus_Macaroni_penguinEudyptes_schlegeli_Royal_penguinOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Paterson_etal_2014a