OUTGROUPUpeneus_taeniopterusUpeneus_asymmetricusUpeneus_crosnieriUpeneus_japonicusUpeneus_luzoniusUpeneus_moluccensisUpeneus_parvusUpeneus_poriUpeneus_quadrilineatusUpeneus_sulphureusUpeneus_tragulaUpeneus_vittatusMullus_argentinaeMullus_barbatusMullus_surmuletusUpeneichthys_lineatusUpeneichthys_porosusUpeneichthys_stottiUpeneichthys_vlamingiiMulloidichtys_flavolineatusMulloidichtys_martinicusMulloidichtys_vanicolensisPseudupeneus_grandisquamisPseudupeneus_prayensisParupeneus_barberinoidesParupeneus_barberinusParupeneus_chrysonemusParupeneus_chgsopleuronParupeneus_ciliatusParupeneus_cyclostomusParupeneus_forsskaliParupeneus_heptacanthusParupeneus_indicusParupeneus_macronemusParupeneus_margaritatusParupeneus_multifasciatusParupeneus_pleurostigmaParupeneus_prophyreusParupeneus_procengenaParupeneus_signatusParupeneus_spilurusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Kim_2002a