Sphenodon_punctatusSimosaurus_gaillardotiOwenetta_kitchingorumAnthodon_serrariusProganochelys_quenstedtiProterochersis_robustaPalaeochersis_talampayensisAustralochelys_africanusKayentachelys_aprixCondorchelys_antiquaHeckerochelys_romaniIndochelys_spatulataMeiolania_platycepsMongolochelys_efremoviHelochelydra_nopcsaiKallokibotion_bajazidiPlatychelys_oberndorferiCaribemys_oxfordiensisNotoemys_laticentralisElseya_dentataChelodina_oblongaPhrynops_geoffroanusErymnochelys_madagascariensisPelomedusa_subrufaPodocnemis_expansaDorsetochelys_delairiPleurosternon_bullockiiGlyptops_plicatulusDinochelys_whiteiNeurankylus_eximiusTrinitichelys_hiattiPlesiobaena_antiquaBoremys_pulchraBaena_arenosaChisternon_undatumPlesiochelys_solodurensisSolnhofia_parsonsiSantanachelys_gaffneyiXinjiangchelys_latimarginalisHangaiemys_hoburensisJudithemys_sukhanoviDracochelys_bicuspisSinemys_lensOrdosemys_leiosToxochelys_latiremysCaretta_carettaChelonia_mydasMesodermochelys_undulatusDermochelys_coriaceaProtochelydra_zangerliMacroclemys_temminckiiChelydra_serpentinaPlatysternon_megacephalumGopherus_polyphemusChrysemys_pictaGeoclemys_hamiltoniiBaptemys_wyomingensisDermatemys_mawiiHoplochelys_crassaStaurotypus_triporcatusSternotherus_odoratusKinosternon_flavescensZangerlia_neimongolensisBasilemys_variolosaAdocus_beatusPeltochelys_durlstonensisApalone_spiniferaLissemys_punctataAnosteira_ornataCarettochelys_insculptaOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Joyce_etal_2011a