DromaeosauridaeArchaeopteryxJeholornisRahonavisShenzhouraptorJixiangornisSapeornisDidactylornisConfuciusornis_sanctusChangchengornisEoconfuciusornisJinzhouornisConfuciusornis_duiChaoyangiaZhongjianornisBoluochiaShenqiornisCathayornisConcornisElsornisEoalulavisEocathayornisEoenantiornisGobipteryxIberomesornisLongipteryxLongirostravisNeuquenornisOtogornisPengornis_houiProtopteryxRapaxavisShanweiniaoVescornisVoronaSchizoouraJianchangornisArchaeorhynchusLiaoningornisSonglingornisAmbiortusLongicrusavisApsaravisHongshanornisYanornisPatagopteryxHollandaYixianornisGansusIchthyornisHesperornisParahesperornisEnaliornisBaptornis_advenusBaptornis_varneriLimenavisVegavisAnasGallusIVPP_V18632IVPP_V18687Eopengornis_martiniOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Hu_etal_2015a