MorganuccodonGobiotheriodonKuehneotheriumTinodonAkidolestesHeishanlestesMaotheriumSpalacolestesSpalacotheridiumSpalacotheriumSpalacotheroidesSymmetrodontoidesSymmetrolestesYaverlestesZhangheotheriumLactodensOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Han_et_Meng_2016a