Geisler, J. H. Uhen, M. D. 2003 Morphological support for a close relationship between hippos and whales Journal of Vertebrate Paleontology 23 991-996 Leptictidae Orycteropus_afer Rattus_norevgicus Vulpavus Canis Arctocyon Eoconodon Hyopsodus Meniscotherium Phenacodus Ambulocetus_natans Artiocetus_clavus Balaenoptera Basilosaurus Delphinapterus_leucas Georgiacetus_vogtlensis Pakicetus Physeter_catodon Protocetus_atavus Remingtoncetus_haurdiensis Rodhocetus Tursiops_truncatus Ankalagon_saurognathus Dissacus_navajovius Dissacus_praenuntius Mongolian_Dissacus Harpagolestes Mesonyx_obtusidens Pachygaena_gigantea Pachygaena_ossifraga Sinonyx_jiashanensis Synoplotherium_canius Hapalodectes_hetangensis Hapalodectes_leptognethus Agriochoerus Amphimeryx Amphirhagatherium_weigelti Bos_taurus Bunomeryx_montanus Cainotherium Camelus Cebochoerus Diacodexis_pakistanensis Wasatchian_Diacodexis Elomeryx_armatus Entelodon Eotylops_reedi Gobiohyus_orientalis Heteromeryx_dispar Hippopotamus_amphibius Homacodon_vagans Hypertragulus Lama Leptomeryx Leptoreodon_marshi Merycoidodon Mixtotherium Odocoileus_virginianus Ovis Perchoerus Poebrotherium Protoceras_celer Raoellidae Sus_scrofa Tayassu_tajacu Tragulus Xiphodon Equus_caballus Mesohippus Heptodon Hyracotherium Hexaprotodon_liberiensis Andrewsarchus Osteology Maximum Parsimony Based on reanalysis of the original matrix. Geisler_et_Uhen_2003a