Proganochelys_quenstedtiRhinochelys_pulchricepsSantanachelys_gaffneyiBouliachelys_suteriNotochelone_costataOcepechelon_bouyaiDesmatochelys_lowiiDesmatochelys_padillaiRhinochelys_nammourensisChelosphargis_advenaCalcarichelys_gemmaProtostega_gigasArchelon_ischyrosDermochelys_coriaceaEosphargis_breineriLepidochelys_olivaceaCaretta_carettaChelonia_mydasEretmochelys_imbricataLepidochelys_kempiiNatator_depressusPuppigerus_camperiArgillochelys_cuneicepsAllopleuron_hofmanniNichollsemys_baieriEochelone_brabanticaCorsochelys_halinchesProcolpochelys_charlestonensisPeritresius_martiniCtenochelys_spCabindachelys_landanensisOligochelone_rupelensisErquelinnesia_gosseletiToxochelys_spSolnhofia_parsoniJurassichelon_oleronensisPortlandemys_mcdowelliPlesiochelys_planicepsPlesiochelys_etalloniPlesiochelys_bigleriSandownia_harrisiBrachyopsemys_tingitanaLeyvachelys_cipadiAngolachelys_mbaxiChelydra_serpentinaMacrochelys_temminckiiSternotherusStaurotypusKinosternon_suburum_hippocrepisDermatemys_mawiiBaptemys_wyomingensisEmarginachelys_cretaceaChrysemys_pictaEmys_orbicularisGeoclemys_hamiltoniiTestudoGopherus_polyphemusChelonoidis_spPlatysternon_megacephalumApalone_spiniferaPelodiscus_sinensisLissemys_punctataCarettochelys_insculptaAllaeochelys_libycaChelus_fimbriatusPhrynops_geoffroanusElseya_dentataChelodinaAraripemys_barretoiPodocnemisPelomedusa_subrufaOrdosemys_sp_IVPP_V12092Judithemys_sukhanoviAnnemys_levensisAnnemys_latiensAnnemys_sp_IVPP_V18106Xinjiangchelys_wusuXinjiangchelys_radiplicatoidesDracochelys_bicuspisKirgizemys_hoburensisKirgizemys_dmitrieviSinemys_gameraSinemys_lensMeiolania_planicepsChubutemys_copelloiEubaena_cephalicaArundelemys_dardeniPleurosternon_bullockiiGlyptops_plicatulusKallokibotion_bajazidiEileanchelys_waldmanniKayentachelys_aprixAustralochelys_africanusAdocus_lineolatusPetrochelys_kyrgyzensisGalianemys_whiteiOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Evers_etal_2019a