LeptictisPhenacodusMeniscotheriumProtolipternaProterotheriumTrigonostylopsAstrapotheriumPyrotheriumNotostylopsHenricosborniaSimpsonotusPleurostylodonThomashuxleyaPampahippusPeriphragnisRyphodonPueliaLeontiniaColpodonAncylocoelusScarittiaHomalodotheriumAsmodeusRhynchippusEurygeniumPascualihippusEomorphippusMorphippusArgyrohippusNesodonAdinotheriumHoffstetteriusPonanskytheriumToxodonColbertiaOldfieldthomasiaCampanorcoAcropithecusUltrapithecusNotopithecusArchaeophylusCochiliusPlagiarthrusProtypotheriumMiocochiliusInteratheriumFedericoanayaTrachytherusPlesiotypotheriumMesotheriumEohyraxPseudhyraxArchaeotypotheriumArchaeohyraxProhegetotheriumHegetotheriumProsotheriumPaedotheriumZalambdalestesMacraucheniaPampahippus_secundusPlexotemnusCoqueniaGriphotherionBoreastylops_Pampatemnus_deuterosPampatemnus_infernalisMartinmiguelia_fernandeziOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Deraco_et_Garcia-Lopez_2016a