LeptictisPhenacodusMeniscotheriumProtolipternaProterotheriumTrigonostylopsAstrapotheriumPyrotheriumNotostylopsHenricosborniaSimpsonotusPleurostylodonThomashuxleyaPampahippusPeriphragnisRyphodonPueliaLeontiniaColpodonAncylocoelusScarittiaHomalodotheriumAsmodeusRhynchippusEurygeniumPascualihippusEomorphippusMorphippusArgyrohippusNesodonAdinotheriumHoffstetteriusPonanskytheriumToxodonColbertiaOldfieldthomasiaCampanorcoAcropithecusUltrapithecusNotopithecusArchaeophylusCochiliusPlagiarthrusProtypotheriumMiocochiliusInteratheriumFedericoanayaTrachytherusPlesiotypotheriumMesotheriumEohyraxPseudhyraxArchaeotypotheriumArchaeohyraxProhegetotheriumHegetotheriumProsotheriumPaedotheriumProticiaColombitheriumBaguatheriumPropyrotheriumCarolozitteliaGryphodonOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Billet_2010a