Protorohippus_venticolumSifrhippus_sandraeOrientolophusCardiolophus_radinskyiHomogalax_protapirinusChowliia_laoshanensisMeridiolophus_expansusIsectolophus_latidensKaragalax_mamikhelensisGandheralophusYimengiaRhodopagus_pygmaeusIndolophusHeptodonHelaletes_nanusDesmatotherium_intermediusParacolodon_fissusDilophodon_minusculusSelenaletes_scopaeusColodon_occidentalisPlesiocolopirus_hancockiHesperaletes_borineyiIrenolophusTelelophus_mediusDeperetellaMinchenoletes_erlianensisSchlosseria_magisterLophialetes_expeditusEoletesAmpholophus_luensisKalakotiaBreviodon_acaresProtapirusTapirusHyrachyus_modestusEphyrachyusFouchiaTriplopus_youjingensisTriplopus_cubitalusEpitriplopus_uintensisTriplopides_rieliProhyracodonHyracodon_nebraskensisArdyniaGobiocerasPappacerasFostercooperiaProeggysodon_qiuiEggysodonJuxiaUrtinotheriumParaceratheriumRostriamynodon_grangeriCaenolophus_promissusAmynodonSharamynodon_mongoliensisCadurcodonParamynodonMetamynodonUintaceras_radinskyiTeletacerasTrigoniasSubhyracodon_occidentalisMenoceras_arikarenseRhinoceros_unicornisOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Bai_etal_2020a