Xenocohippus_osborniPachynolophus_livinierensisDanjiangia_pingiLambdotherium_popoagicumEotitanopscf_EotitanopsPalaeosyopsEotitanops_pakistanensisBalochititanops_haqiBunobrontops_savegeiMesatirhinus_juniusDesmatotitan_tukhumensisAcrotitan_ulanshirehensisDolichorhinus_hyognathusSphenocoelus_uintensisMicrotitan_mongoliensisFossendorhinus_diploconusMetarhinus_paterMetarhinus_fluviatilisMetarhinus_abbottiSthenodectes_incisivumTelmatherium_validusWickia_brevirhinusMetatelmatherium_ultimumXylotitan_cenosusQufutitan_zhouiEpimanteoceras_formosusProtitan_grangeriProtitan_minorRhinotitan_kaiseniRhinotitan_andrewsiNanotitanops_shanghuanensisPollyosbornia_altidensPygmaetitan_panxianensisGnathotitan_berkeyiAktautitan_hippopotamopusBrach_transylvanicumMetatitan_primusMetatitan_relictusMetatitan_khaitshinusNasamplus_progressusProtembolotherium_efremoviEmbolotherium_andrewsiEmbolotherium_grangeriPachytitan_ajaxProtitanotherium_emarginatumDiplacodon_elatusDiplacodon_giganParabrontops_gobiensisEubrontotherium_clarnoensisParvicornus_occidentalisProtitanops_curryiNotiotitanops_mississippiensisDianotitan_lunanensisDuchesneodus_uintensisMegacerops_coloradensisMegacerops_kuwakatarhinusMaobrontops_paganusOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Averianov_etal_2018b