Tapirus_terrestris_outgroupHyrachyus_eximius_outgroupTrigonias_osborni_outgroupRonzotherium_filholi_outgroupAceratherium_incisivumAlicornops_simorrenseBrachypotherium_brachypusBugthirhinus_praecursorChilotherium_anderssoniDiaceratherium_aginenseDiceratherium_armatumDicerorhinus_sumatrensisDiceros_bicornisGaindatherium_browniHispanotherium_beonenseLartetotherium_sansanienseMenoceras_arikarenseMesaceratherium_gaimersheimenseMesaceratherium_paulhiacenseMesaceratherium_welcommi_sp_novPlesiaceratherium_mirallesiPleuroceros_blanfordiPleuroceros_pleurocerosProsantorhinus_douvilleiProtaceratherium_albigenseProtaceratherium_minutumRhinoceros_unicornisSubhyracodon_occidentalisOsteologyMaximum ParsimonyBased on reanalysis of the original matrix.Antoine_etal_2010a