(Cardiolophus_radinskyi,(Hallensia_matthesi,(Pliolophus_vulpiceps,(Hyracotherium_leporinum,(Hyracotherium_levei,(Hyracotherium_remyi,(Orolophus_maldani,((Propalaeotherium_gaudryi,Propalaeotherium_sudrei,Propalaeotherium_cf_gaudryi_AUM,(Propalaeotherium_hassiacum,(Eurohippus_parvulus,(Lophiotherium_pygmaeum,Lophiotherium_cervulum,Lophiotherium_sp_AUM))),(Propalaeotherium_isselanum,(Leptolophus_nouleti,(Palaeotherium_magnum,Plagiolophus_minor)))),((Pachynolophus_duvali,Pachynolophus_sp_AUM),(Pachynolophus_eulaliensis,Pachynolophus_ruscassierensis_n_sp,(Pachynolophus_cesserasicus,Pachynolophus_livinierensis)))))))))));