(Sphenodon_punctatus,(Owenetta_kitchingorum,Anthodon_serrarius,(Odontochelys_semitestacea,(Proganochelys_quenstedti,((Proterochersis_robusta,(Kayentachelys_aprix,Condorchelys_antiqua,Heckerochelys_romani,Eileanchelys_waldmani,Indochelys_spatulata,Niolamia_argentina,Ninjemys_oweni,Warkalania_carinaminor,Meiolania_platyceps,Chubutemys_copelloi,Peligrochelys,Kallokibotion_bajazidi,Spoochelys_ormondea,Chengyuchelys,((Dorsetochelys_delairi,Neurankylus_eximius,Solnhofia_parsonsi,Thalassemys_moseri,Santanachelys_gaffneyi,(Patagoniaemys_gasparinae,Otwayemys_cunicularius),(Simosaurus_gaillardoti,((Notoemys_laticentralis,(Platychelys_oberndorferi,Caribemys_oxfordiensis)),(Prochelidella_cerrobarcinae,Phrynops_geoffroanus,Yaminuechelys_maior,Araripemys_barretoi,(Elseya_dentata,Myuchelys_latisternum),(Chelus_fimbriatus,(Chelodina_colliei,Chelodina_longicollis)),(Pelomedusa_subrufa,(Erymnochelys_madagascariensis,Podocnemis_expansa))))),(Pleurosternon_bullockii,(Glyptops_plicatulus,Dinochelys_whitei)),(Trinitichelys_hiatti,Plesiobaena_antiqua,(Boremys_pulchra,Baena_arenosa,Chisternon_undatum)),(Portlandemys_macdowelli,Plesiochelys_etalloni),((Toxochelys_latiremys,((Caretta_caretta,Chelonia_mydas),(Mesodermochelys_undulatus,Dermochelys_coriacea))),((Macroclemys_schmidti,Macroclemys_temminckii,Protochelydra_zangerli,Chelydra_serpentina,Platysternon_megacephalum),(Emarginachelys_cretacea,(((Emys_orbicularis,(Shachemys_laosiana,(Mongolemys_elegans,(Echmatemys_wyomingensis,(Geoclemys_hamiltonii,((Stylemys_nebraskensis,(Eurotestudo_hermanni,(Gopherus_polyphemus,(Chelonoidis_gringorum,Chelonoidis_chilensis)))),(Chrysemys_picta,Trachemys_scripta))))))),((Baptemys_wyomingensis,Dermatemys_mawii),(Hoplochelys_crassa,(Xenochelys_formosa,Staurotypus_triporcatus,(Sternotherus_odoratus,Kinosternon_flavescens))))),(((Basilemys_variolosa,Adocus_beatus),(Yehguia_tatsuensis,Basilochelys_macrobios)),(Anosteira_ornata,Carettochelys_insculpta,(Apalone_spinifera,Plastomenus_aff_thomassii,Pelodiscus_sinensis,Lissemys_punctata))))))),(Siamochelys_peninsularis,(Xinjiangchelys_wusu,Annemys_levensis,Annemys_latiens,Xinjiangchelys_radiplicatoides,Xinjiangchelys_junggarensis),(Hangaiemys_hoburensis,Changmachelys_bohlini,Judithemys_sukhanovi,Dracochelys_bicuspis,Ordosemys_skull,Manchurochelys_manchoukuoensis,Xiaochelys_ningchengensis,(Ordosemys_leios,Liaochelys_jianchangensis),(Sinemys_brevispinus,Sinemys_lens,Sinemys_gamera))),(Helochelydra_nopcsai,Naomichelys_speciosa)),(Sichuanchelys_chowi,(Mongolochelys_efremovi,Sichuanchelys_palatodentata))))),(Palaeochersis_talampayensis,Australochelys_africanus))))));