(Parahippus_leonensis,(M_gunteri,(M_primus,(Astrohippus,Pliohippus,M_carrizoensis,M_stylodontus,M_intermontanus,(Hippidion,Onohippidium,Equus,Dinohippus),(Protohippus,Calippus),(Merychippus_isonesus,M_sejunctus),((M_coloradense,(Pseudhipparion,Neohipparion)),(M_insignis,(Hipparion,(M_goorisi,(Cormohipparion,Nannippus)))))))));