(Outgroup,(Liostrea_delta,Liostrea_eduliformis,Pernostrea,(UNC_13497b,((Lopha_haidingeriana,Dendostrea_frons,Lopha_solitaria,(Rastellwn_macropterwn,(Rastellwn_carinatum,(Agerostrea_falcata,Agerostrea_mesenterica)))),(Nicaisolopha_nicasei,(Ostreola_conchaphila,((Saccostrea_palmula,Striostrea_marganitacea),(Crassostrea_columbiensis,Crassostrea_virginica)))))),(Liostrea_hisingeri,(Praexogyra_acuminata,Praexogyra_calceola)),(Nanogyra_nana,((Amphidonte,Ceratostreon_texanum,Ceratostreon_zueatherfordensis),(Exogyra_ponderosa,Exogyra_laeviscula_slash_tigrina,Ilymatogyra_arietina))),(Gryphaea_arcuataeformis,Gyphaea_arcuata,(Gryphaea_sportella,(Gryphaea_pictavensis,Catinula_beaumonti),(Gryphaea_nebrascensis,(Texigraphaea_roemeri,(Pycnodonte_roanokensis,Parahyotissa_mcgynti)))))));