(Simpsonotus,(Pleurostylodon,Colbertia,(Notopithecus,Antepithecus,((Antofagastia,(Punapithecus,(Johnbell,Ignigena,(Eopachyrucos,(Santiagorothia,Proargyrohyrax,(Archaeophylus,Federicoanaya,Brucemacfaddenia,(Plagiarthus,(Cochilius,Interatherium)),(Protypotherium,Miocochilius,Progaleopithecus))))))),(Transpithecus,Guilielmoscottia)))));