(Ancestor,((Hyrachyus_eximius,Triplopus_cubitalis,(Rhodopagus_radinskyi,Siamolophus_krabiense)),(Heptodon,(Helaletes_nanus,((Desmatotherium_intermedium,Dilophodon),((Colodon_occidentalis,Colodon_inceptus,(Irdinolophus_mongoliensis,(Irdinolophus_tuiensis,Teleolophus))),(Plesiocolopirus,Protapirus)))))));