(INDOTESTUDO,((HERMANNI,BOETTGERI),((RUSTAMOVI,(HORSFIELDII,KAZACHSTANICA)),(((TERRESTRIS,HIGHLAND),(FLOWERI,ANTAKYENSIS)),(IBERA,((ARMENIACA,(ZAGROS,(NIKOLSKII,(ZARUDNYI,(ANAMURENSIS,(MARGINATA,WEISSINGERI)))))),((GRAECA,NABEULENSIS),(SOUSSENSIS,(LIBYA,(KLEINMANNI,NEGEV))))))))));