(Tapirus_terrestris_outgroup,(Hyrachyus_eximius_outgroup,(Trigonias_osborni_outgroup,(Ronzotherium_filholi_outgroup,(((Protaceratherium_minutum,(Plesiaceratherium_mirallesi,(((Aceratherium_incisivum,Alicornops_simorrense),(Diaceratherium_aginense,(Brachypotherium_brachypus,Prosantorhinus_douvillei))),((Dicerorhinus_sumatrensis,(Diceros_bicornis,Rhinoceros_unicornis)),(Gaindatherium_browni,Lartetotherium_sansaniense))))),((Mesaceratherium_paulhiacense,(Mesaceratherium_gaimersheimense,Protaceratherium_albigense)),(Pleuroceros_blanfordi,Pleuroceros_pleuroceros))),(Subhyracodon_occidentalis,(Diceratherium_armatum,(Menoceras_arikarense,(Hispanotherium_beonense,(Bugthirhinus_praecursor,(Chilotherium_anderssoni,Mesaceratherium_welcommi_sp_nov)))))))))));