(Dromaeosauridae,(Archaeopteryx,(Jeholornis,(((Jinguofortis,Chongmingia),(Sapeornis,((Protopteryx,((Monoenantiornis,((Fortunguavis,(Eoenantiornis,(Linyiornis,((Gretcheniao,(((Boluochia,Longipteryx),(Shanweiniao,(Longirostravis,Rapaxavis))),(Gobipteryx,Vescornis))),((Piscivorenantiornis,(Mirusavis,(Shangyang,(Eocathayornis,(((Neuquenornis,(Concornis,Qiliania)),(Eoalulavis,Cathayornis)),(Elsornis,Cruralispennia)))))),(Dunhuangia,Pterygornis)))))),(Shenqiornis,(Sulcavis,(Longusunguis,(Zhouornis,(Bohaiornis,Parabohaiornis))))))),((Pengornis,Eopengornis),(Chiappeavis,Parapengornis)))),(Archaeorhynchus,(Jianchangornis,(((Schizooura,Mengciusornis),(Bellulornis,((Vorona,Patagopteryx,Eogranivora),(Apsaravis,(Xinghaiornis,Dingavis))))),((Songlingornis,((Piscivoravis,(Yixianornis,(Iteravis,(Gansus,((Ichthyornis,(Vegavis,(Anas,Gallus))),(Enaliornis,(Baptornis_advenus,(Baptornis_varneri,(Hesperornis,Parahesperornis))))))))),(Similiyanornis,(Yanornis,Abitusavis)))),(Hongshanornis,(Parahongshanornis,(Longicrusavis,(Archaeornithura,Tianyuornis))))))))))),(Eoconfuciusornis_zhengi,(Confuciusornis_sanctus,Confuciusornis_dui,(Changchengornis_hengdaoziensis,Yangavis)))))));