(A_dydimacantha,((C_cf_pellizari,(C_acutifrons,((((S_aculeata,(S_granulosa,S_tarimuensis)),((S_whittingtoni,(S_crossi,(S_analoga,S_lacrima))),(S_angulata,S_gadwanensis))),(C_minutula,((C_gaoluoensis,(C_pentagonalis,(K_matchensis,((((K_minaretta,(L_ludvigseni,(L_exopthalma,L_hintzei))),(K_acuticaudata,K_lacrimosa)),((C_carrerai,(T_bituberculata,(C_gracielae,C_sanjuanina))),(K_piojensis,(K_salasae,K_sterrenae)))),(C_riojana,K_forbesi))))),(C_pusilla,C_salopiensis)))),(C_alata,C_changshanensis)))),((C_fabiani,((C_gonioloba,K_typicalis),(C_sulcatus,A_grindrodi))),(C_rushtoni,(C_tauwhena,C_wrighti)))));