(Hesperocyoninae,(Borophaginae,(Leptocyon,((Vulpes,(Urocyon,Otocyon)),((q_Pseudalopex_q,((Pseudalopex,Dusicyon),(Lycalopex,Chrysocyon,((Cerdocyon,Nyctereutes),(Atelocynus,Speothos))))),(Canis,(Cuon,Lycaon)))))));