(Pleurostylodon_modicus,(Pampahippus_arenalesi,(Plexotemnus_complicatissimus,Puelia_coarctatus),((Eomorphippus_obscurus,((Rhynchippus_equinus,Rhynchippus_pumilus,Rhynchippus_brasiliensis,(Pascualihippus_boliviensis,(Nesodon_imbricatus,Proadinotherium_leptognathum,Adinotherium_ovinum)),(Argyrohippus_praecox,Argyrohippus_fraterculus)),(Eurygenium_pacegnum,Eurygenium_latirostris))),(Leontinia_gaudryi,Scarrittia_canquelensis))));