(Cardiolophus_radinskyi,(Hallensia_matthesi,(Pliolophus_vulpiceps,(Hyracotherium_leporinum,(((Propalaeotherium_hassiacum,((Propachynolophus_gaudryi,Propalaeotherium_sudrei),(Propalaeotherium_isselanum,(Leptolophus_nouleti,(Palaeotherium_magnum,Plagiolophus_minor))))),(Eurohippus,(Lophiotherium_pygmaeum,Lophiotherium_cervulum))),((Propachynolophus_levei,(Propachynolophus_maldani,Propachynolophus_remyi)),(Pachynolophus_eulaliensis,(Pachynolophus_cesserasicus,(Pachynolophus_duvali,Pachynolophus_livinierensis)))))))));