(Outgroup,(Preondactylus,((Dimorphodon,Peteinosaurus),((Versperopterus,(Jeholopterus,(Batrachognathus,(Anurognathus,Dendrorhynchoides)))),((Raeticodactylus,(Austriadactylus,(Campylognathoides,Eudimorphodon))),(Fenghuangopterus,((Sordes,(Scaphognathus,(Pterorhynchus,(Dorygnathus,(Nesodactylus,Rhamphorhynchus))))),(Changchengopterus,(Cacibupteryx,(Darwinopterus,((((Pterodactylus,Cycnorhamphus),(Cathayopterus,(Eosipterus,(Ctenochasma,(Gegepterus,(Pterodaustro,(P_longicolum,Gnathosaurus))))))),((G_rhamphastinus,(G_cristatus,(Noripterus,(Dsungaripterus,Tatal_pterosaur)))),(((Lonchodectes,(Tupuxuara,Thalassodromeus)),((Shenzhoupterus,Chaoyangopterus),(Azhdarcho,(Quetzalcoatlus,Zhejiangopterus)))),(Tupandactylus,(Tapejara,(Sinopterus,Huaxiapterus)))))),((((((Cearadactylus,Guidraco),(Ludodactylus,(Ornithocheirus,(Anhanguera,Coloborhynchus)))),(Pteranodon,(Nyctosaurus,Muzquizopteryx))),(Haopterus,(Istiodactylus,Nurhachius))),(Boreopterus,Zhenyuanopterus)),((Moganopterus,(Elanodactylus,(Feilongus,Beipiaopterus))),(Gladocephaloideus,Pterofiltrus))))))))))))));