(Outgroup,(Preondactylus,((Dimorphodon,Peteinosaurus),((Dendrorhynchoides,(Jeholopterus,(Anurognathus,Batrachognathus))),(((Austriadactylus,Raeticodactylus),(((Scaphognathus,Sordes),(Pterorhynchus,((Cacibupteryx,Fenghuangopterus),(Dorygnathus,(Nesodactylus,Rhamphorhynchus))))),(Darwinopterus,(Changchengopterus,((Cycnorhamphus,(Pterodactylus,(Cathayopterus,(((Gegepterus,(Pterodaustro,(Eosipterus,Ctenochasma))),(Pterodactylus_longicolum,Gnathosaurus)),(((Moganopterus,Feilongus),(Beipiaopterus,Elanodactylus)),(Gladocephaloideus,Pterofiltrus)))))),(((((Ludodactylus,((Cearadactylus,Guidraco),(Ornithocheirus,(Anhanguera,Coloborhynchus)))),(Haopterus,(Istiodactylus,Nurhachius))),(Pteranodon,(Nyctosaurus,Muzquizopteryx))),(Boreopterus,Zhenyuanopterus)),((Germanodactylus_rhamphastinus,((Germanodactylus_cristatus,Noripterus),(Dsungaripterus,Tatal_pterosaur))),((((Lonchodectes,(Tupuxuara,Thalassodromeus)),(Azhdarcho,(Quetzalcoatlus,Zhejiangopterus))),(Shenzhoupterus,Chaoyangopterus)),(Sinopterus,(Huaxiapterus,(Tapejara,Tupandactylus))))))))))),(Campylognathoides,Eudimorphodon))))));