(Boreosomus,(Birgeria,(Chondrosteus,(Peipiaosteus,((Polyodon,Protopsephurus),(Psammorhynchus,(Scaphirhynchus,((Acipenser_oxyrinchus,Acipenser_transmontanus,((Acipenser_baeri,(Huso_huso,Acipenser_ruthenus)),(Acipenser_stellatus,(Pseudoscaphirhynchus_kauffmani,Pseudoscaphirhynchus_hermanni)))),(Acipenser_brevirostrum,Acipenser_fulvescens)))))))));