(Proganochelys_quenstedti,(Australochelys_africanus,((Chubutemys_copelloi,(Meiolania_planiceps,(Kallokibotion_bajazidi,((((((Toxochelys_sp,(Corsochelys_halinches,((Ocepechelon_bouyai,(((((Rhinochelys_pulchriceps,Rhinochelys_nammourensis),(Santanachelys_gaffneyi,Notochelone_costata)),(Bouliachelys_suteri,Chelosphargis_advena)),(Desmatochelys_lowii,Desmatochelys_padillai)),(Calcarichelys_gemma,(Protostega_gigas,Archelon_ischyros)))),((Dermochelys_coriacea,Eosphargis_breineri),(Nichollsemys_baieri,((Allopleuron_hofmanni,((Argillochelys_cuneiceps,(Chelonia_mydas,(Natator_depressus,(Eretmochelys_imbricata,(Lepidochelys_kempii,(Lepidochelys_olivacea,Caretta_caretta)))))),(Procolpochelys_charlestonensis,((Puppigerus_camperi,Eochelone_brabantica),(Oligochelone_rupelensis,Erquelinnesia_gosseleti))))),(Cabindachelys_landanensis,(Peritresius_martini,Ctenochelys_sp)))))))),((Solnhofia_parsoni,((Plesiochelys_planiceps,(Jurassichelon_oleronensis,Portlandemys_mcdowelli)),(Plesiochelys_etalloni,Plesiochelys_bigleri))),(Leyvachelys_cipadi,(Brachyopsemys_tingitana,(Sandownia_harrisi,Angolachelys_mbaxi))))),(((Chelonoidis_sp,(Platysternon_megacephalum,(Chelydra_serpentina,Macrochelys_temminckii))),((Dermatemys_mawii,(Baptemys_wyomingensis,(Emarginachelys_cretacea,((Kinosternon_suburum_hippocrepis,(Sternotherus,Staurotypus)),(Adocus_lineolatus,(((Apalone_spinifera,Petrochelys_kyrgyzensis),(Pelodiscus_sinensis,Lissemys_punctata)),(Carettochelys_insculpta,Allaeochelys_libyca))))))),(Chrysemys_picta,((Emys_orbicularis,Geoclemys_hamiltonii),(Testudo,Gopherus_polyphemus))))),(Araripemys_barretoi,((Elseya_dentata,(Phrynops_geoffroanus,(Chelus_fimbriatus,Chelodina))),(Pelomedusa_subrufa,(Podocnemis,Galianemys_whitei)))))),(((Ordosemys_sp_IVPP_V12092,Dracochelys_bicuspis),(Sinemys_gamera,Sinemys_lens)),(Judithemys_sukhanovi,(Kirgizemys_hoburensis,Kirgizemys_dmitrievi)))),(Annemys_levensis,(Annemys_latiens,(Annemys_sp_IVPP_V18106,(Xinjiangchelys_wusu,Xinjiangchelys_radiplicatoides))))),(Arundelemys_dardeni,(Eubaena_cephalica,(Pleurosternon_bullockii,Glyptops_plicatulus))))))),(Eileanchelys_waldmanni,Kayentachelys_aprix))));