(Gorilla_gorilla,(Pan_troglodytes,(Ardipithecus_ramidus,(Australopithecus_anamensis,(Sahelanthropus_tchadensis,(Australopithecus_afarensis,(Australopithecus_garhi,((Australopithecus_africanus,(Australopithecus_sediba,((Homo_habilis,(Kenyanthropus_platyops,Homo_rudolfensis)),(small_bodied_Dmanisi,((Homo_antecessor,(Homo_ergaster,((Homo_heidelbergensis,Homo_neanderthalensis),(Homo_sapiens,D4500)))),(Homo_erectus,Homo_floresiensis)))))),(Paranthropus_aethiopicus,(Paranthropus_robustus,Paranthropus_boisei))))))))));