(Archaeopteryx,(Jeholornis,((Sapeornis,((Protopteryx,((shenqiornis,(Orienantius,((((Junornis,Gretcheniao),((Zhouornis,(Eoenantiornis,(CUGB_P1202,((Concornis,(Eoalulavis,Cathayornis)),((Iberomesornis,(Eocathayornis,Gobipteryx)),(Neuquenornis,Sulcavis)))))),(Longusunguis,(Bohaiornis,(Qiliania,Parabohaiornis))))),((Elsornis,Longipteryx),(Shanweiniao,(Longirostravis,Rapaxavis)))),(Vescornis,(Pengornis,Fortunguavis))))),(Eopengornis,Parapengornis))),((Schizooura,Bellulornis),((Jianchangornis,(((Longicrusavis,(Apsaravis,Hongshanornis)),((Gansus_yumenensis,Gansus_zheni),(Ichthyornis,((Hesperornis,Parahesperornis),(Vegavis,(Anas,Gallus)))))),(Yanornis,Yixianornis))),(Archaeorhynchus,Patagopteryx))))),(Eoconfuciusornis,(Changchengornis,(Confuciusornis,Vorona))))));