(Cardinolophus,(Homogalax,((Protomoropus,Pappomoropus,(Paleomoropus,(Lophiaspis,((Litolophus,Grangeria_canina),(Grangeria_anarsius,(Eomoropus_pawnyunti,((Eomoropus_ulterior,Eomoropus_amarorum),(Eomoropus_quadridentatus,(Eomoropus_minimus,(SDM_84006,(Schizoherium_avitum,(Moropus_elatus,Ansiodon_grande)))))))))))),(Lophiodon,(Hyrachyus_eximus,Heptodon_calciculus)))));